Arachnologische Mitteilungen 35

spider fauna of semi-dry grasslands 55 Most of the species are eurytopic, only 13 are ste- notopic. Of these, Agalenatea redii , Cheiracanthium virescens , Enoplognatha latimana , Xysticus acerbus and Zelotes longipes are stenotopic to dry grassland and sandy habitats. Hypsosinga pygmaea is stenoto- pic to heathlands. All stenotopic species occur only with a low number of specimens (Tab. 2). Considering the number of investigation days and traps the largest number of specimens could be caught at sites D2 and D3, followed by D1, while the number of specimens of sites D4 – D7 is low (Tab. 3). The highest number of species is given at D1 followed by D4 in contrast to D2 and D7 with the lowest number of species. Regarding the Brillouin-Index D5 and D4 show the highest diversity whereas the values for D2 and D3 are noticeable low. Thus in comparison with all other sites (E B = 0.66 – 0.72), the Brillouin-Eveness of site D2 (E B = 0.48) and D3 (E B = 0.46) is also low while the largest value for Brillouin-Eveness could be calculated for D5 (E B = 0.79). On the basis of the PCA it is possible to separate four species groups (Fig. 2). In the plot the species composition of the sites D1, D2 and D3 [A] is clearly distinguished from that of the sites D4 and D5 [B] and D6 and D7 [C]. The first axis may reflect a humidity gradient with dry habitats on the left and humid sites on the right. Along the second axis probably a vegetation cover gradient is given. The PCA shows an overlap between the sites D1, D2 and D3. Thus the first group of spiders occur on sites which show typical conditions of dry grassland, rough meadows and grazing: sparse and low-growing vegetation with places of bare sand. The community comprises 22 species. Some of them are exclusively found at these sites (e.g. Centromerita concinna , Drassyllus lutetianus ), others seem to prefer these structures (e.g. Alopecosa accen- tuata , Pardosa agrestis , P. palustris , Xerolycosa mini- ata ). Only one species, Xysticus acerbus , is stenotopic to this habitat. Most of the species seem to be able to cope with a wide ecological amplitude and thus family/species RL HB D1 D2 D3 D4 D5 D6 D7 A D M Clubiona pallidula * e . . . . 1 . . 1 0,01 pt Clubiona reclusa * e 1 . . . 1 . . 2 0,02 pt Clubiona terrestris * e . . . . . . 2 2 0,02 pt Corinnidae Phrurolithus festivus * e 15 1 1 2 2 2 . 23 0,21 pt Gnaphosidae Drassodes pubescens * e 9 5 9 6 10 4 1 44 0,39 pt Drassyllus lutetianus * e 1 . 16 . . . . 17 0,15 pt Drassyllus pusillus * e 42 39 75 62 78 3 7 306 2,73 pt Haplodrassus signifer * e 38 30 49 17 15 6 17 172 1,54 pt Haplodrassus umbratilis * e 2 . . . . . . 2 0,02 pt Micaria pulicaria V e 2 . . . 1 . . 3 0,03 pt Micaria subopaca R s3 . . . . . 1 . 1 0,01 pt Zelotes electus V e 2 . 2 30 7 . 1 42 0,38 pt Zelotes latreillei * e 8 . 12 25 23 3 . 71 0,63 pt, hs Zelotes longipes V s1 . . 2 . . . . 2 0,02 pt Zelotes petrensis * e 16 . 3 3 13 . . 35 0,31 pt Zelotes subterraneus * e 1 . . . . 4 . 5 0,04 pt Zoridae Zora silvestris * s4 3 . . . . . . 3 0,03 pt Zora spinimana * e 1 . . . . 1 1 3 0,03 pt Philodromidae Philodromus cespitum * e . . . 1 . 1 . 2 0,02 pt, hs Philodromus praedatus * s3 . . . . . . 1 1 0,01 pt Thomisidae Xysticus acerbus 3 s1 3 5 6 . 1 . . 15 0,13 pt Xysticus audax * e 2 . . . . . . 2 0,02 hs Xysticus cristatus * e 15 11 25 10 6 3 13 83 0,74 pt, hs Xysticus kochi * e 7 12 8 3 2 2 6 40 0,36 pt, hs Salticidae Euophrys herbigrada 2 e . . . 2 . 1 . 3 0,03 pt Evarcha falcata * e 1 . . . . . . 1 0,01 pt Heliophanus flavipes * e . . . 7 . 1 . 8 0,07 hs Phlegra fasciata V e 4 1 2 2 3 . . 12 0,11 pt, hs Talavera aequipes V e 1 . 1 . . . . 2 0,02 pt ∑ 2162 2725 3275 986 922 542 583 11195 82 49 62 70 58 60 46