Arachnologische Mitteilungen 54

60 P. Bezděčka, K. Bezděčková & T. Kvamme Results and discussion Newly collected in Norway We collected altogether 163 specimens of harvestmen from 21 species, 12 genera and three families in the studied sites. Five species are newly reported from Norway. Family Nemastomatidae Mitostoma chrysomelas (Hermann, 1804) L–09: 2 (( ; L–12: 2 )) , 1 ( (all under stones). Nemastoma bimaculatum (Fabricius, 1775) L–03: 2 (( ; L–12: 2 )) (all under stones). Nemastoma lugubre (O. F. Müller, 1776) L–13: 2 (( (under a stone). Family Phalangiidae Lacinius dentiger (C. L. Koch, 1847) L–12: 1 ) , 1 ) (on a tree), a new record for Norway. Lacinius ephippiatus (C. L. Koch, 1835) L–03: 2 )) (under a stone). Lacinius horridus (Panzer, 1794) L–13: 4 )) , 1 ( (on a rock and on soil), a new record for Norway. Lophopilio palpinalis (Herbst, 1799) L–12: 2 )) , 2 (( (under vegetation); L–13: 3 )) (under a stone). Mitopus morio (Fabricius, 1779) L–12: 2 )) (on vegetation); L–13: 2 )) , 3 (( (on soil). Oligolophus hanseni (Kraepelin, 1896) L–3: 1 ) , 1 ( ; L–04: 1 ) ; L–12: 1 ) , 1 ( (all under stones); L–20: 1 ) (on a wall). Oligolophus tridens (C. L. Koch, 1836) L–12: 3 )) , 3 (( ; L–15: 3 )) (all under stones), L–20: 1 ) (on a tree). Opilio canestrinii (Thorell, 1876) L–02: 2 )) ; L–08: 2 )) ; L–18: 1 ) ; L–20: 2 )) , 1 ( (all on walls). Opilio parietinus (De Geer, 1778) L–17: 2 )) (on tree); L–20: 1 ) (on a wall). Opilio saxatilis C. L. Koch, 1839 L–20: 2 )) (on a wall), a new record for Norway. Paroligolophus agrestis (Meade, 1855) L–02: 4 (( (under wood); L–03: 2 (( ; L–04: 2 )) , 4 (( ; L–12: 5 )) , 2 (( ; L–13: 1 ) , 4 (( ; L–15: 2 )) , 2 (( (all under stones); L–20: 3 (( (on a tree). Phalangium opilio Linnaeus, 1758 L–01: 2 )) (on vegetati- on in a garden); L–02: 1 ( (on a wall); L–03: 1 ) , 1 ( (on a tree); L–04: 2 (( (on a rock); L–05: 1 ) ; L–06: 2 (( (on vegetation); L–09: 2 (( (on a wall); L–12: 3 (( ; L–13: 2 (( ; L–15: 1 ) , 1 ( ; L–16: 2 )) (all on vegetation); L–20: 1 ) , 1 ( (on a wall). Rilaena triangularis (Herbst, 1799) L–16: 2 juv. (under vege- tation). Family Sclerosomatidae Leiobunum blackwalli Meade, 1861 L–01: 1 ) , 2 (( (on a wall), a new record for Norway. Leiobunum gracile Thorell, 1876 L–07: 2 )) (on a rock); L–09: 1 ) , 1 ( ; L–10: 1 ) , 1 ( ; L–11: 1 ) ; L–13: 1 ( ; L–15: 1 ) , 1 ( ; L–20: 2 (( (all on trees). Leiobunum limbatum L. Koch, 1861 L–08: 2 juv.; L–19: 2 (( ; L–20: 1 ) (all on walls), a new record for Norway. Leiobunum rotundum (Latreille, 1798) L–01: 1 ) , 1 ( ; L–02: 2 (( ; L–08: l ) , 1 ( (all on walls); L–09: 2 )) , 2 (( ; L–10: 2 (( ; L–13: 2 )) , 2 (( ; L–14: 1 ) , 1 ( ; L–15: 1 ) , 2 (( ; L–20: 1 ( (all on trees). Nelima gothica Lohmander, 1945 L–10: 1 ) , 1 ( (on a tree). In total 21 species were recorded, out of which five species were reported from Norway for the first time. The species L . horridus and O. saxatilis had already been found in all Nordic mainland countries. L. dentiger had been recorded in Finland, L. blackwalli in Sweden and Denmark. The presence of L. limbatum is presently known only from Norway. We found this species in the cities of Kristiansand and Oslo, in the vi- cinity of harbours.This might suggest an introduction of this species to Norway by ship or other types of transportation. Additionally, we assume that several other harvestmen species may be reported from Norway in the near future, e.g. Dicranopalpus ramosus (Simon, 1909), Odiellus spinosus (Bosc, 1792), the still unnamed species Leiobunum sp. “A” (sensu Wijnhoven et al. 2007) or Nelima semproni Szalay, 1951. The revised harvestmen checklist of the Nordic countries In the past ten years, the list of harvestmen from Nordic countries has undergone a number of changes. The species currently known from the individual countries are summa- rized in Tab. 1. New records and changing distribution patterns of Opi- liones species have been reported in Northern Europe (e.g. Gederaas et al. 2012, Uddström et al. 2013, 2016b, Enghoff et al. 2014, Fritzén et al. 2015).The list presented here is cer- tainly not definitive. Future research will undoubtedly bring a number of new faunistic records and may also reveal the occurrence of additional species in Nordic countries. Two years of first description differ from those given by Stol (2007): Trogulus tricarinatus Linnaeus, 1767 (instead of 1758 – it was formerly wrongly assigned to the 10 th instead of the 12 th edition of Linnaeus’ ‘Systema naturae’) and Paroli- golophus meadii O. Pickard-Cambridge, 1891 (corrected from 1890, according to Sykes 1941). According to the revision of Martens & Schönhofer (2016) all specimens of the formerly recorded Leiobunum ru- pestre (Herbst, 1799) from Scandinavia belong to L. gracile , a senior synonym of Leiobunum tisciae Avram, 1968. Leiobunum limbatum was reported for Sweden by Stol (2007) based on Martens (1978).However, according to Jons- son (2017 pers. comm.), there is neither older voucher mate- rial nor new findings supporting the presence of this species in Sweden. Specimens of Platybunus bucephalus (C. L. Koch, 1835) recorded from Finland were questioned by Stol (2007). La- ter the species was deleted from the Finish list – all revised specimens were juvenile specimens of Rilaena triangularis (cf. Uddström et al. 2013, Fritzén et al. 2015). Information con- cerning the presence of Opilio dinaricus and Mitostoma chryso- melas in Finland is based on Uddström & Rinne (2016b).The authors stated that it is quite probable that these species were imported into Finland with raw wood and it is not certain whether they could actually establish in Finland. Acknowledgements We are very grateful to Professor Henrik Enghoff, University of Copenhagen (Denmark),Dr. Lars J. Jonsson, Kristianstad University, Kristianstad (Sweden) and Dr. Annika Uddström, Finnish Environ- ment Institute,Helsinki (Finland) for providing valuable information.

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