Arachnologische Mitteilungen 55
28 A. Šestáková, A. Mock, J. Christophoryová & P. Gajdoš or Asthenargus . Until now few males of the species have been identified, but still not formally described. However, Blick et al. (in prep.) are in process of publishing a comprehensive ta- xonomy of P. aenigmaticus and its relatives, thus we refrain from describing the male in the present paper. Habitat. This species seems to prefer caves and similar habi- tats (Blick & Kreuels 2002), given that most of the recorded specimens have been discovered there (Blick & Kreuels 2002, Růžička & Buchar 2008, Růžička et al. 2013). Slovak speci- mens were also found in a cave and 75 cm beneath the sur- face in the granitoid mesovoid shallow substratum. Further findings refer to post-mining dumps (Staręga 1996), railway underpasses (Kůrka et al. 2015), beech forests (Milasowszky et al. 2015), and oak and fir forests (Rozwałka & Olbrycht 2017).They have also been identified in different types of ag- ricultural landscape, including fields, open grasslands, mea- dows, fallow lands, shrubland and urban and rural landscapes such as parks, orchards and abandoned mining and quarry sites (Blick & Kreuels 2002). Colonisation of new habitats presumably occurs via ballooning (Blandenier & Fürst 1998). Status . This species is red-listed in Central Europe due to very rare and scarce findings; endangered in Poland (EN) (Staręga et al. 2002), vulnerable in Germany (VU) (Blick et al. 2016) and listed under least concern in the Czech Republic (LC) (Řezáč et al. 2015). Distribution. European species recorded in Austria, Belgi- um, the Czech Republic, Germany, Great Britain, Italy, Lu- xemburg, Poland and Switzerland (Blick & Kreuels 2002, Helsdingen 2013, World Spider Catalog 2017). Acknowledgements We are grateful to Vlastimil Růžička and Theo Blick for help in determining the spiders and their helpful comments on an earlier version of the manuscript. We thank Peter Ľuptáčik for the photos of Plavecká Cave, Raymond Marshall for English correction and the Editor and three reviewers for their constructive suggestions. This study was partly funded by grant projects Nos. 2/0171/16 and 1/0199/14 from the Slovak Grant Agency VEGA. References Blandenier G & Fürst P-A 1998 Ballooning spiders caught by a suc- tion trap in an agricultural landscape in Switzerland. In: Selden PA (ed.) Proceedings of the 17th European Colloquium of Arach nology,Edinburgh 1997.British Arachnological Society,Burnham Beeches, Bucks. pp. 177-186 Fig. 7: Pseudomaro aenigmaticus female from the Jajcajov jarok Valley, Slovakia. A. dorsal view; B. ventral view; C. epigyne Fig. 8: Pseudomaro aenigmaticus male from the Plavecká Cave, Slovakia. A. dorsal view; B. ventral view
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