Arachnologische Mitteilungen 57

Checklist of the spiders of Left-Bank Ukraine 61 We marked with three exclamation marks (!!!) species registe­ red for the first time in Ukraine, with (!!) species registered for the first time in Left-Bank Ukraine, and with (!) species cited in literature as new to Left-Bank Ukraine in 2017, i.e. after the issue of the Catalogue and Addendum I (Polchaninova & Prokopenko 2013, 2017). We marked boundaries of the species’ geographic ranges by a letter in case they are running through the study area: N – northern, NW – north-western, W – western, NE – north-eastern, E – eastern, S – southern, SE – south-eastern, and SW – south-western, intr – introdu­ ced. The list is based on original and literature-derived data that can be checked on the available collection materials. If a species was recorded from material in a lost collection (or lost individuals from available collections) and its finding had not been proved later by other authors, we referred it to the category “need confirmation”. A list of such species is gi­ ven separately after the main species list. We also compiled a list of erroneous records based on misidentifications. Seven species were mentioned in both “erroneous” and “doubtful” lists since their misidentifications were proved for some lo­ calities, but remained unconfirmed for the others (Table 2, 3, electronic appendix). The rationale for the exclusion of these species from the main list is given in the Catalogue and Ad­ dendum I (Polchaninova & Prokopenko 2013, 2017). Species distribution is given based on Mikhailov (2013), Polchaninova & Prokopenko (2013), Kovblyuk & Kastrygina (2015), Ponomarev (2017), Ponomarev et al. (2017), Nentwig et al. (2018), WSC (2018) and the authors’ personal data. Results To date, 741 spider species from 270 genera and 34 families have been recorded from Left-Bank Ukraine. The number of families has decreased compared with the previous list, since Argyroneta, the only regional genus in Cybaeidae, was transferred to the family Dictynidae. Thus, Cybaeidae was excluded from the araneofauna of Left-Bank Ukraine. The following genera are the most speciose: Pardosa – 20 species, Clubiona , Xysticus – 17 species each, Gnaphosa , Zelotes, Wal­ ckenaeria – 16 species, Alopecosa – 15 species, Agyneta – 14 species, Micaria – 11 species , and Haplodrassus, Centrome­ rus – 10 species. Four species listed here ( Alopecosa steppica , Palludiphantes pillichi , Calositticus inexpectus , Pellenes allegrii ) are new to the Ukrainian fauna; two species ( Gnaphosa opaca and Centromerus abditus ) are new to Left-Bank Ukraine. Four species ( Altella hungarica , Erigonoplus spinifemuralis , Walcken­ aeria wunderlichi , Synageles ramitus ) were recorded as new to Ukraine in 2017 (Ponomarev et al. 2017) after the publication of the Addendum I of the catalogue, and are now added to the general list. Ostearius melanopygius and Heliophanus kochii were absent in the 2013 catalogue, Gnaphosa rufula and Dys­ dera longirostris were restored to the list from the doubtful records, and Gnaphosa dolanskyi was distinguished from G. modestior and described as new to science (Řezáč et al. 2018). Thus, the previous list has been replenished with 15 species. On the other hand, G. modestior and G. moesta have been excluded, so the list of erroneous records now comprises 41 species while that of doubtful records accounts for 47 species. The family Linyphiidae is the most species-rich in Left- Bank Ukraine (215 species, 29% of the fauna), with Gnapho­ sidae and Salticidae following it (83 and 79 species, respec­ tively, about 11% each). The ratio of families changes from north to south across zones and subzones (Fig. 3). The pro­ portion of Linyphiidae declines from 33% in the forest zone to 16% in the bunchgrass steppe subzone (steppe II). In con­ trast, the ratio of Gnaphosidae grows from 7% in the forest zone to 13% in the steppe subzones. Fluctuation of the relati­ ve species richness of other families is not so evident. Diversi­ ty of major spider families is the most even in the bunchgrass steppe subzone (steppe II) due to the lower ratio of Linyphi­ idae (Fig. 3). The sagebrush-bunchgrass steppe subzone has not been analyzed here because of the lack of sufficient data on its spider fauna. The richest is the araneofauna of the steppe I (583 spe­ cies). All major families, except Linyphiidae, are the most di­ verse there (Table 1). The steppe II is the poorest (375 spider Fig. 1: Map of Ukraine with the outlined study area Fig. 2: Borders of natural zones and subzones in Left-Bank Ukraine. Zones: F – forest, F-St – forest-steppe, St – steppe; subzones: St I –forb-bunchgrass steppes, St II – bunchgrass steppes, St III – sagebrush-bunchgrass steppes